PSU Nittany Lions

2012 record: 8-4
2012 conference record: 6-2 (second, Leaders Division)
Returning starters: Offense: 8; defense: 6, kicker/punter: 2

Top returners

RB Zach Zwinak, WR Allen Robinson, TE Kyle Carter, OG John Urschel, OT Donovan Smith, DE Deion Barnes, LB Glenn Carson, CB Adrian Amos

Key losses

QB Matt McGloin, FB Michael Zordich, C Matt Stankiewitch, DT Jordan Hill, LB Michael Mauti, LB Gerald Hodges, CB Stephon Morris

2012 statistical leaders (*returners)

Rushing: Zach Zwinak* (1,000 yards)
Receiving: Allen Robinson* (1,013 yards)
Tackles: Gerald Hodges (109)
Sacks: Deion Barnes* (6)
Interceptions: Michael Mauti (3)

Spring answers

1. "Tight End U." If there's one position the Nittany Lions don't have to worry about, it's this one -- and that's probably why some PSU players have taken to dubbing the university "TEU." Kyle Carter's injured wrist should be just fine once the season rolls around, and there's plenty of depth here. Teammates have pointed to the offseason work of 6-foot-7 target Jesse James, who really came on strong in the second half of last season. He was also the receiving star in the annual spring scrimmage with five catches and 77 yards. Couple him with Matt Lehman, Brent Wilkerson and Adam Breneman, and it wouldn't be a surprise to see quite a few formations with multiple tight ends on the field.

2. Versatility at a premium. Bill O'Brien doesn't mind moving players around. Wideouts Malik Golden and Trevor Williams switched over to the secondary, and Williams has especially picked up the position quickly. But those two aren't the only to play at different positions. Adrian Amos can play safety or cornerback, and a lot of Penn State's younger DBs have the ability to slide between those two. Penn State's trying to combat a lack of depth with versatile players here, so players who can play at multiple spots are especially valuable.

3. Young standouts. Several true and redshirt freshmen could contribute heavily this season, and O'Brien has praised multiple first-year players for picking things up quickly. On defense, DT Austin Johnson looks to be a starter after a redshirt season, and LB Nyeem Wartman has a leg up on an injured Ben Kline. On offense, WR Eugene Lewis made a one-handed grab in the spring scrimmage to show he can make the tough catches, and RB Akeel Lynch has also made a strong case for playing time. PSU doesn't historically have many four-year starters, but this year could change that.

Fall questions
1. Quarterback question marks. Neither option, early enrollee Tyler Ferguson nor incoming freshman Christian Hackenberg, has ever thrown a pass in the FBS -- and one of those two players will be the Penn State starter. Inexperience is a big concern, and the QB will have to learn a complicated offense in a short period of time. Hackenberg has a lot of potential and Ferguson showed glimpses, however inconsistent, in the spring game. But the offense's strength last season was the quick no-huddle offense -- and it remains to be seen whether either of these signal callers can pull the fast playing style off.

2. Withstanding lack of depth. O'Brien has gotten this team down to about 67 scholarships in preparation for 2014 when the 65-scholarship limit kicks in, so depth is a real concern this season. If a quarterback or linebacker becomes injured, PSU could be in trouble. The Lions need to remain healthy to have a shot at repeating last year's success. And one injury could really have a ripple effect on this team. Health is one question, one uncertainty, that can't be answered anytime soon.

3. Kicking game. Sam Ficken was just 14-of-21 on field goals last season and didn't make a single kick over 39 yards. He did wind up converting his last 10 attempts, but his inconsistency carried over in the spring game when he missed a 37-yard field goal and an extra point. O'Brien was known for leaving the special-teams unit on the sideline a lot on fourth downs last season and, if Ficken struggles again, that would put even more pressure on the young quarterbacks. Or force O'Brien to use incoming walk-on kicker Chris Gulla.

STATE COLLEGE, Pa. -- Akeel Lynch clapped his hands as he walked off the field Saturday. He smiled, waved to a few cameras and seemed happier than some of the fans who snagged autographs from PSU coaches.

This was the tailback's debut, the first time he competed in front of more than a scattered high school crowd. Last season he redshirted and watched his teammates from Beaver Stadium's sideline. So, on a chilly afternoon during the Blue-White Game, with grass stains covering his white pants, Lynch tried to take it all in.

That smile wasn't just because of his numbers -- 13 carries, 83 yards, one touchdown -- or because fans applauded as he jogged toward the tunnel. He has been waiting for this for a long time, and when asked about it he couldn't help but grin again.

"I was just happy because I finally played in Beaver Stadium," he said. "This is something I dreamed about since I got that Penn State jacket as an 8-year-old, and actually coming out and playing in Beaver Stadium -- finally getting a grass stain on my jersey -- it's a dream come true. I was all smiles."

That jacket, a blue $20 varsity jacket purchased by his mom in Erie, Pa., came during fifth or sixth grade, before Lynch ever heard of the school. He liked the color blue, so his mom thought he'd like the garb. And, when he moved from Canada to Buffalo as a high school junior, his mom found that same jacket and gave it back.

"Wouldn't it be funny if you go to Penn State?" he remembered her asking.

His mom was there in the crowd Saturday, undoubtedly smiling along with her son. After all, Lynch was the name that rested on most people's minds immediately after the game. His teammates, fellow tailback Zach Zwinak and linebacker Glenn Carson, talked in the weeks leading up to the Blue-White Game about his progress.

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STATE COLLEGE, Pa. -- Saturday's annual scrimmage, known as the Blue-White Game, will offer fans a nice reprieve as they wait more than 20 weeks for the college season to kick off.

Saturday will be the first time most of last season's freshmen will play in front of a crowd, the first time fans can size up the quarterback race and the first time the media can see the progress this team has made over the spring.

The Blue-White weekend has taken on a carnival-type atmosphere these past few years, and there's plenty to see. But on the field, NittanyNation takes a closer look on what fans should keep an especially close eye on.

QUARTERBACK RACE

Let's get the obvious out of the way. Steven Bench threw just eight passes last season, and the media has seen little of Tyler Ferguson. Both quarterbacks have been lauded for their ability to scramble -- Stephon Morris said he probably wouldn't even label Bench as a pocket passer -- but both are basically a mystery. Can Bench guide this offense? Will Ferguson outshine him? Saturday's scrimmage is far from the be-all, end-all, but it is a start to answering some of those questions.

Bill O'Brien said in the past no quarterback separated himself yet. Maybe, just maybe, someone will gain an edge Saturday.

BRING ON THE MAN COVERAGE

Defensive coordinator John Butler acknowledged last season that PSU couldn't play be as aggressive in the secondary because of the depth. But that is improved this season. Jordan Smith and Anthony Smith enrolled early, while wideouts Malik Golden and Trevor Williams switched to defensive back.

PSU began practicing the nickel this spring, and fans can expect finally to see that package this season. There's no telling who might start alongside Adrian Amos come August -- Jordan Lucas is currently practicing with the first team -- and fans should keep an eye on the young corners here.

PROJECTED (RS) FRESHMAN STARTERS

DT Austin Johnson and LB Nyeem Wartman are just redshirt freshmen, but it already looks as if they'll crack the starting lineup this season. Both very well could wind up as the rare four-year PSU starter, and expectations are high for these two.

Johnson already is up to 302 pounds, and O'Brien has praised his ability since he was asked about his top freshmen last season. And the hard-hitting Wartman, whom PSU fans already are familiar with, blocked a punt in his PSU debut before an injury in Week 2 that sidelined him for the season (and allowed him to pick up a medical redshirt). With the departures of Mike Mauti and Gerald Hodges, Wartman will have to play well right off the bat for PSU to remain strong here. One recruit said he was especially impressed watching No. 5, because he was all over the field during one practice. Let's see what they can do in a scrimmage.

(Also, Akeel Lynch and Eugene Lewis might not be starters ... but is there anyone who doesn't plan to keep a close eye on them?)

HOW MUCH BETTER HAVE THE BEST GOTTEN?

Practice observers and teammates have pointed constantly to Allen Robinson when asked who has impressed so far this spring. He broke the single-season school record for receptions last season, and he has gained needed weight while maintaining his speed. Robinson was the best in the Big Ten last year, and now he's even better. That's hard to picture on the field.

Mike Hull, Deion Barnes, Zach Zwinak, Amos, etc. all have earned a lot of praise this spring. Zwinak has improved his strength, Barnes is shoring up his run-stopping, Hull is embracing a starting role ... and Amos? Well, he's probably PSU's most versatile player. Returner, safety, cornerback -- he can do everything. And it'll be interesting to see just how much he does Saturday.

TIGHT END U?

It's pretty incredible just how much this position has evolved in about 15 months. Kyle Carter won't play in the Blue-White Game, but fans still will be able to look at Matt Lehman, Jesse James and Brent Wilkerson.

MLB Glenn Carson mentioned James as the player who has impressed him the most overall. He broke out during Carter's absence late in the year, and he boasts good speed for a 6-foot-7 target. James has "red-zone target" written all over him, and it'll be interesting to see how this young corps does in the scrimmage. It'll be an even bigger bonus if Adam Breneman is able to play.

STATE COLLEGE, Pa. -- Adrian Amos was known as a shy, do-everything DB last season. He was a man of many highlights and few words -- but Wednesday afternoon was a little different.

The junior stood tall, carried a smirk and had a lot to say: He's looking forward to playing more nickel, the secondary looks a lot better, man coverage has greatly improved, Trevor Williams is making a nice switch to defense, etc. But, despite the roll Amos was on, PSU's versatile DB had to stop and stare at the ground a few moments when asked about himself.

What position don't you play in the secondary?

Amos hesitated slightly, seemingly going over the positions in his mind. "There's -- there's not one," he said.

What position are you best at?

"Psssh ... I don't know," he added. "People say my natural position is safety, but then people say I'm better at corner. But I feel comfortable playing all over."

That's good news for Penn State, because the staff isn't finished trying him out elsewhere. The 6-foot All-Big Ten honorable mention has not only practiced at both safety spots, focused on the boundary corner, played some field corner -- but he discovered just this week he's also the team's nickelback.

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STATE COLLEGE, Pa. -- Every spring, a few players step up and offer the first glimpses of an upcoming breakout season. Often, the media can only guess and project since they're not at every practice, weightlifting session and team meeting.

The players, however, are there and can see who might be the next Deion Barnes or Allen Robinson. So, NittanyNation polled several players inside the Lasch Football Building on Wednesday and also took information from Wednesday teleconferences.

The question: Who's made the biggest impact this spring or who's been the biggest surprise? Here's what Penn State's veterans had to say:

Linebacker Glenn Carson: "I'm really impressed with Jesse James. He's had a great winter. That kid's just an animal. I'm really excited to watch him play this year. I think he's a freak athlete. I think he can do amazing things. And if you haven't been impressed by him yet, I think you're going to be even more impressed by him next year."

Offensive guard John Urschel: "Ty Howle is a great center and a great football player. I'd say he's one of the most underrated football players on our team. You don't hear about him enough. I don't think you guys talk about him enough. I don't think we're going to have any problems with him playing at center."

Defensive tackle DaQuan Jones: "I really think Austin Johnson, Derek Dowrey and Brian Gaia. They're young, but they're showing a lot of ability. They're going to be great down the road. I'm happy seeing their competitiveness in spring ball."

Safety Malcolm Willis: "It's a number of guys. If I had to pick one, it'd probably be John Urschel. He comes in every day early and gets his workout in early. And pound for pound, he's probably one of the strongest guys on the team. Mike Hull, he's another guy who works his butt off. Adrian Amos. Zach Zwinak -- I work out with him in the weight room every day, and the amount of weight he can put up, it's crazy. Those are a couple of the guys who have really impressed me so far."

Offensive tackle Adam Gress: "I'd have to say a lot of the defensive linemen are coming along. All of the younger guys are really working hard, and the guys I was playing against on the scout team last year, now those guys are players. They're people we compete with. So that group's made a lot of progress. ... Austin Johnson's definitely made a lot of progress."

Linebacker Mike Hull: "Nyeem [Wartman] has looked good so far. He's doing a great job filling in. I think he's going to be a great player for us. We just need to all come together as a unit. He's looking good so far, so we just have to see how he progresses."

Defensive end Deion Barnes: "I think it would be Kyle Baublitz. I think he's doing a great job right now. Whoever's been talking to him has done a great job because he's come out of his shell. I've never seen him like this before."

STATE COLLEGE, Pa. -- Bill O'Brien wasn't overly pleased with his quarterbacks at the beginning of Wednesday's practice. The fiery coach yelled early on at the four signal-callers, "You do not want me to play quarterback, because I can't throw."

The media were allowed to attend about 30 minutes of Penn State's fifth spring practice, and the defense seemed to get the better of the offense during a brief battle. Juco quarterback Tyler Ferguson took snaps with the the first team during the offense-defense competition, while Steven Bench went with the second team. (They alternated first-team snaps during walk-throughs.)

The offense had four chances to reach a first down, and Ferguson opened up by connecting with Allen Robinson on a seven-yard pass. That was the offensive highlight of the two series, however, as linebacker Glenn Carson stopped Zach Zwinak at the line of scrimmage, then Ferguson was nearly "sacked" on a play-action pass.

Bench didn't fare any better and didn't complete a pass in four downs. In what might have been the best overall play, defensive end Brad Bars made nice penetration, then leaped in the air to deflect a screen pass. His teammates yelled, "Oooooh," because of the impressive vert, but Bench actually caught the rebound. Still, the play was whistled dead.

Bill Belton ran twice without much success, and Bench then held on to the ball for what seemed like an eternity. At least two linemen could've sacked him had he not been wearing a red shirt, and he ended up scrambling to the outside. That play also was blown dead.

The offense was forced to do five hit-its as a result of losing the competition. Obviously, eight snaps and a handful of passes don't genuinely reflect the quarterbacks' overall success, but it at least offered a glimpse.

Injury report: Tight end-turned-offensive tackle Garry Gilliam did not take part in practice because of an injury to his right calf. He's week-to-week. ... Stephen Obeng-Agyapong (shoulder) and Kyle Carter (wrist) wore red shirts but practiced. ... DT DaQuan Jones spent the first few minutes of practice with the first team during walk-throughs but was not wearing pads and did not participate in contact drills. The nature of his injury was not immediately clear, but a spokesman said it wasn't serious. ... Tight end Adam Breneman also wore a red shirt -- something he didn't do in the first practice -- and had his right arm taped up.

Visitors: Members of the Cornell coaching staff were on hand to watch practice, and two 2014 recruits also walked around the practice field to take in the sights.

ESPN Watch List QB Michael O'Connor (Bradenton, Fla./IMG Academy) and cornerback Marcus Whitfield talked to their respective position coaches and watched from the sideline.

Starting lineup: At wide receiver, Robinson and Brandon Moseby-Felder lined up as the starters, while both Austin Johnson and C.J. Olaniyan practiced with the first-team defense.

In a curious move, Donovan Smith was limited while Eric Shrive replaced him. Smith received some one-on-one instruction at times, and it wasn't clear if he was injured.

Regardless, it didn't seem serious. Smith is the clear starter, and reporters were told this was the first practice where Smith was treated like that.

Special teams' woes: The field-goal unit practiced what appeared to be extra points. (The yardage markers run parallel to the posts, so it's not clear.) On one snap, center Ty Howle tossed it back a little low and the holder wasn't able to get a handle it.

It rocketed behind the kicker when Ryan Keiser couldn't grab it in time. PSU was inconsistent on special teams last season, and mistakes like that are exactly what it can't afford in the regular season.

STATE COLLEGE, Pa. -- While Bill O'Brien's voice carried over the field Monday afternoon, quarterbacks coach Charlie Fisher directed his players in a calmer manner.

During position drills at Penn State's first spring practice, Fisher stood about five yards in front of his four quarterbacks as they took turns taking three- and five-step drops. The quartet of red jerseys would look at Fisher, quickly scan the field and then throw to a stationary receiver.

"Eyes here," Fisher told sophomore Steven Bench in a conversational tone, pointing to his his right. "Work through it, work through it."

Bench or Tyler Ferguson could wind up as the starting quarterback come Aug. 31, and Monday offered a glimpse of the two signal-callers. Media were invited to attend 20 minutes of open practice, and O'Brien began by calling together a competition: A defensive back would line up against a wideout or tight end in press coverage, while Bench and Ferguson would alternate snaps.

The first team -- offense or defense -- to win three battles would be declared the winner. The losers would be forced to perform five hit-its. It was more for honor than anything and only four passes were thrown while the quarterbacks tried to shake off the rust.

Bench began by just overthrowing Allen Robinson on a roughly 35-yard pass, and Ferguson then hit Jesse James in stride downfield on an over-the-shoulder grab. Bench followed that up by throwing behind his target on cross route, and Ferguson barely overthrew Matt Lehman for two straight incompletions.

The offense, along with Ferguson and Bench, then hit the turf for their hit-its while the defense cheered.

"They're both athletic, they both can throw the football," O'Brien said during a Monday news conference. "Now it's going to depend on how well they make decisions and how accurately they throw the ball.

"They sit in the front row, they pay attention, they take a lot of notes. It's a fun group to be around."

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STATE COLLEGE, Pa. -- Montee Ball set the NCAA record for total touchdowns Saturday, but the Badgers were unable to improve their conference record as Penn State rallied late for the 24-21 overtime win.

Sam Ficken connected on a 37-yard field goal to open up overtime. Wisconsin responded by fumbling and nearly tossing an interception before Kyle French missed a field-goal attempt wide left from 45 yards.

Wisconsin took the early lead in the game by scoring two quick touchdowns -- including Ball's record 79th touchdown -- but Ted Roof's defense adjusted and allowed just seven points the rest of the way. Matt McGloin, who had only 47 passing yards at halftime, rebounded in the second half and paced the offense.

With the win, Penn State's season ends at 8-4 (6-2 Big Ten) while Wisconsin drops to 7-5 (4-4).

It was over when: French missed a 45-yard field goal attempt wide left in overtime. Glenn Carson nearly picked off a pass on third down, and Sean Stanley pushed Wisconsin back after forcing a fumble.

Game ball goes to: Zach Zwinak. He set career highs in carries (36) and rushing yards (179) and was the constant that Penn State's offense needed. He was never tackled for a loss and even outrushed Ball by 68 yards.

Stat of the game: 27 -- the number of yards Ball rushed for in the second half. He averaged 6.5 yards per carry in the first half but only mustered 1.9 yards a carry in the second half. Defensive tackle Jordan Hill helped clog up the middle to limit Ball's production.

Turning point: Early in the final quarter with PSU trailing 14-13, Bill O'Brien found himself on Wisconsin's 41-yard line. He opted to go for it, and McGloin connected with Jesse James on a crossing route that ended up as a 41-yard touchdown. That gave PSU its first lead.

Unsung hero: Hill. He's not yet 100 percent, but he might have had his best game of his career Saturday. He finished with 12 tackles and a pair of sacks. He was a big reasons PSU was able to shut down Ball in the second half.